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After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise ...

The beloved Galápagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Tortoises can live for more than 100 years in the wild and close to double that in captivity. This tortoise was born in her native habitat likely in the 1880s.

But now Gramma, a Galápagos tortoise, has eaten her last cactus fruit, the San Diego Zoo reported. She arrived in San Diego ...

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Gramma the tortoise, believed to be 141 years old and the oldest resident of the San Diego Zoo, was euthanized after suffering from progressive bone deterioration because of her age.

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