The beloved Galapagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

But now Gramma, a Galápagos tortoise, has eaten her last cactus fruit, the San Diego Zoo reported. She arrived in San Diego ...

A Jacksonville woman's quarter-sized tortoise grew into something unexpected: a conservation movement. Now she's using her ...

National History, a son of Maxfield, and Devon Island, by Practical Joke, lead an uncoupled Godolphin entry into a 1600m maiden at Meydan.