Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that ...

Tortoises can live for more than 100 years in the wild and close to double that in captivity. This tortoise was born in her native habitat likely in the 1880s.

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galpagos tortoise ...

But now Gramma, a Galápagos tortoise, has eaten her last cactus fruit, the San Diego Zoo reported. She arrived in San Diego ...

How the solitary leopard tortoise survives in the heart of the wild.

Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died at an estimated 141 years of age.